Ptilodactyla

Ptilodactyla toe-winged beetle. Urbana, Illinois, USA

Cytilus alternatus

Cytilus alternatus - pill beetle. Pennsylvania, USA.

Lutrochus

Travertine beetles live in streams with travertine limestone. Arizona, USA.

Lutrochus

The long claws of these travertine beetles help them grip the substrate under running water.

Heterocerus

Variegated mud-loving beetle. Quincy, California, USA.